Inter-annual and spatial climatic variability have led to a balance between local fluctuating selection and wide-range directional selection in a perennial grass species. - Keep, T; Rouet, S; Blanco-Pastor, J L; Barre, P; Ruttink, T; Dehmer, K J; Hegarty, M; Ledauphin, T; Litrico, I; Muylle, H; Roldán-Ruiz, I; Surault, F; et al. (Annals of Botany, 2021)